The organism is what we call a psychotroph. It will grow very, very slowly at refrigerator temperatures, so four degrees is not a problem, and it will get to very, very high numbers over a period of 59 days, which is the shelf life of a cooked, cured meat product in a vacuum package. It does not need oxygen. It will grow without oxygen.
It's relatively pH insensitive. It will grow at very high pHs--pH 9, pH 4.5. That's not a problem for it. And it will also grow at body temperature, and we know that, because it kills people.